martes, 29 de noviembre de 2016

Has any species of animal become extinct in the past 20 years?

Sadly, many. So many that scientists are talking about a massive extinction induced by humans, the sixth in the history of life:
Accelerated modern human-induced species losses: Entering the sixth mass extinction

The last animals added are:
2015 — Eastern Cougar, Puma Concolor Couguar
2013 – Formosan Clouded Leopard, Neofelis nebulosa brachyura
2012 — Pinta Island Tortoise, Chelonoidis abingdoni
2011 — Vietnamese Rhino, Rhinoceros sondaicus annamiticus


2009 — Christmas Island Pipistrelle, Pipistrellus murrayi
2007 — Chinese Paddlefish, Psephurus gladius
2007 — Yangtze River Dolphin, Lipotes vexillifer
etc, etc.
If you want to visit this sad list, here is the link:
Here's Every Single Animal That Became Extinct In The Last 100 Years
As a meta-analysis, a study published by experts from Stanford, Princeton and the University of  California-Berkeley declared the world’s vertebrates are going extinct  114 times faster than the natural rate of extinction.

domingo, 27 de noviembre de 2016

If birds are dinosaurs, then why aren't all vertebrates fishes? Are they only dinosaurs when speaking cladistic, or also Linnaean?


 I understand that, with clades, birds are dinosaurs, but when considering clades, birds are fishes, cnidarians, et cetera. This distinction is next to useless for the general purposes of language, where calling everything a bacteria would be meaningless. Thus, for the purposes of casual language, are birds really dinosaurs?

Another question of Quora, if you read these entries you already know that I considered the site interesting before they chose quantity (traffic) instead of quality. They also have a politic of top writers (those who attract more readings) that has made havoc with the level of answers related to animals and ethology.
However, I once wrote on Quora this text. Enjoy...
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Birds are dinosaurs for both systems of classification.
Birds are dinosaurs from both viewpoints, although it carries some problems for Linnean classification (1). And don't missunderstand me, I do really like Linnean classification and prefer that it be the first one to be taught to children because it is a great description of the world as we see it, such as Cladistics can't be by definition, because they have to include features of extinct organisms. Here in Spain children are introduced to classification overly early, in the first grades of Primary, and how could they understand the links between reptiles and birds that Cladistics bring when they are under 10 years?.
Coming back to dinosaurs, it is a group that was not defined on the range of characters we can observe on a living animal that include external features such as appearance, skin traits (having fur, feather, glands, layers...), metabolism traits (such as weter they could maintain a level of internal temperature independently of enviromental conditions or not, chambers in the heart, etc), skeletal morphology and others. It wasn't possible because the description was made from the fosil remains that only depicted skeletal features (2), hence all the group was typed from some few relevant skeletal traits. There is not way of comparing how they descripted mammals, birds and reptiles with the way they did with dinosaurs.
What happened is that accordingly with that key skeletal features, birds must be included into the dinosaur group (from a Linnean viewpoint) or are in the dinosaur clade (from a Cladistic one). Birds fulfill all the "must to" to be named dinosaurs by their own and not as derived from dinosaurs. Of course that there is a lot of polemics sorrounding this point, for some anatomist birds display very original adaptations to flight that would tell them apart from more "primitive" dinosaurs (3). Some even maintain that they could derive from an earliest ancestor that was only related to dinosaurs, but was not a dinosaur (4). I think that the scientits that defend those viewpoints have failed to demonstrate that birds lack the skeletal characters that make them dinosaurs, so far at least; in fact the critics I link bellow are debunked. Probably we will see more of this polemic in the following years, and this is the kind of discussions I like, because in spite that they can be really boring for taxonomists, for general public show some really interesting features of the animal studied. And also, it is like a real scientific soap opera on the internet.
But for the purpose of casual language, birds can be called birds, and we can forget for a while that they are dinosaurs.
Finally, in our dayly life, as far as we are not interested in the evolutionary history of some organism or some feature in some organism, I think it is more useful, and amazing, talking about them in the way we see and experience our present world, and here is where Linnean classification makes the best role. Do you see those beautiful long legged horses as fishes?. Indeed, nobody does. The insistence of calling birds dinosaurs come from the fact that the polemics between supporters and the remaining detractors of birds as dinos still continues, nothing is more noisy than a battle field.
__ Links and an introduction to some problems and controversies derived from dinosaurs classification. I only recomend point 1 and 4 to dino freaks.
(1) Well, this point is annoyingly long, I agree. But as I have seen so many people confused with the idea that dinosaurs are birds, I like to explain why they feel so stranged with it.
The problems that Linnean classification must face due to that birds are actually dinosaurs derive from that dinosaurs include animals that highly resemble reptiles and others that, well, are birds.
For the lay person the problem can be post as "So when we think about the term dinosaur, what are we thinking about a bird or a reptile, or both?". In fact the matter reminds me the Necker cube , an optical illusion in what we can see the same drawing of a cube looking up, down or both alternately.
Regarding dinosaurs, there is not such dilema, as I have told in the text, dinosaurs were defined on different basis than any living animal Class (reptiles and birds are Classes accordingly to Linnean classification), so there is not contradiction in that dino groups animals from both Classes. It is a little uncomfortable for Linnean supporters, but not a nuclear misile that will destroy the concept.
Cladistics supporters, on the other hand, have more strong feelings about the Linnean Class reptilia (i.e reptiles). Some schools of thought would like to redefine it in a way that includes birds. But they also have to face their own problems, because it is not clear how to group the different extant types of reptiles accordingly to evolutionary characters. For the moment reptiles are a nightmare for evolutionary biologists and cladistics zealots, good for the old brave scaled animals. Who wants that their children are told that birds an reptiles are the same at the tender age of seven?. I don't.
In fact dinosaurs are not the only group that defy the traditional Linnean grouping, just the most popular. If Synapsid would have been so know as dinos, we would have assisted to a long and vively discussion about wether they are retiles or mammals or what. Finally synapsid were split from reptiles.
I have brought about the example of synapsid to talk about the complexity of classifying extinct animals that depict characters that make them close to two different Linnean Classes, species or whatever. In the end, making classifications is drawing borders that looked from an aerial perspective seem to be very clear, but the closer you get to these borders, the more artifitial and debatable they appear. This is the lesson that we, lay people, can draw from the problems with classifying dinosaurs.
(2) What is the scientific diagnosis of what is a dinosaur, and what is just another archosaur? Several skeletal characteristics are currently used as diagnostic dinosaurian features. You may also view a large-screen picture of a dinosaur skeleton for a lesson in anatomy : Morphology of the Dinosauria
(3) Are birds dinosaurs? New evidence muddies the picture (bear in mind that the objections exposed in this article are nowadays debunked).
(4) Longisquama (debunked. But I really find Longisquasma amazing, in spite that I only see the head half of an animal skeleton over some feathered thing).

sábado, 26 de noviembre de 2016

What are some amazing moments of bonding between animals from different species?


I love this question on Quora because I love animals. I find it uplifting, so let's have a nice moment. Enjoy...
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1)  Mzee is a Aldabra giant tortoise (Aldabrachelys gigantea), a species which habitat is in the Seychelles. Owen is exactly what you could expect from a hipoppotamus.

 Let's start with one of the most surprising pals I had never expected. Because one of them is a reptil, and the other a mammal. You may think that in this case the bond could only flow in one direction (from mammal to reptil), very probably, but it was fundamental for the well being of the young mammal.
Owen and Mzee


They live in the Haller Park in Kenya. In December 2004, tthe terrible tusnami separated Owen, then a juvenile hippo, from his herd. he was rescued and carried to the Haller Park Rescue Center. Having no other hippos to interact with, Owen immediately attempted to  bond with Mzee, whose large domed shell and brown color resembled an  adult hippo. They formed an odd friendship that continued until 2006,  when it was determined that Owen had grown too large to safely interact  with Mzee. Today they live in separated enclosures, and Owen is doing a great job in socializing with other hippos, which reveals that the effect of the perceived bonding with the tortoise was very positive for the young mammal.


2) Whales interacting with dolphins: Sperm Whales 'adopt' a deformed dolphin, and whales giving dolphins a lift.

These kind of bondings can occur in the wild. Despite they can be temporal, we can't deny that they defy our understanding of humans as the only ones that can understand another species's animal suffering and help them, and also...what puzzles me more...that they enjoy playing with another species.

Let's start by an amazing example, this video depicts a very rare interaction between sperm whales and an  adult bottlenose dolphin with a spinal malformation (i.e. scoliosis).  This represents the first time this type of non-agonistic (friendly)  interaction has been recorded for sperm whales. We published a  description of these interactions in the scientific journal "Aquatic  Mammals".



Surprising, isn't it?. But what if I tell you that scientists have filmed whales playing with dolphins...playing, not harrasing or bullying.

Whales give dolphins a lift.





Many species interact in the wild, most often as predator and prey. But  recent encounters between humpback whales and bottlenose dolphins reveal  a playful side to interspecies interaction. In two different locations  in Hawaii, scientists watched as dolphins "rode" the heads of whales:  the whales lifted the dolphins up and out of the water, and then the  dolphins slid back down. The two species seemed to cooperate in the  activity, and neither displayed signs of aggression or distress.

3)  Adopted pets can form bonds incredible strong, because they know what is not to have them. When there is a disability included in the equation, the bondings are truly impresionant.

I've talked about wild animals in a rescue center, and wild animals in the wild, but the greatest source of these unthinkable bondings are just average people homes

Pwditat the rescued straycat and Terfel the blind dog:


 And, on the other hand, Murdock the blind adopted cat and the family dog who adopted him as his best friend (the name of the dog wasn't mentioned in the article).


Here are the links to know more about them.
http://www.lifewithcats.tv/2012/...
http://lovemeow.com/2013/10/murd...

4) There is an entire channel of NatGeo dedicated to tell this kind of stories.
 http://channel.nationalgeographi...
Here is the story of a ten y.o hen and a very handicapped toy dog, that are best buddies:

martes, 22 de noviembre de 2016

Why are poop-throwing chimpanzees more intelligent?

 http://phys.org/news/2011-11-poop-throwing-chimps-intelligence.html

Ok, some questions on Quora are incredibly funny. And, ah, oh, yes...some field studies too bizarre.
This is one of the first things I wrote on Quora, be patient with my English, I was starting, and also forgive me some typos...but poo throwing chimps and intelligence, is there anything more exciting?.
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From suchanaughtymonkey.wordpress.com

Are there -not handycapped- chimps that can't poop throw ?.
The paper says that better poop throwers were more intellegent becaus:

After making their  discovery regarding the parts of the brain that appear to be involved in  better throwing in chimps, the team tested the chimps and found that  those that could throw better also appeared to be better communicators  within their group, giving credence to their idea that speech and  throwing are related. Interestingly, they also found that the better  throwing chimps didn’t appear to posses any more physical prowess than  other chimps, which the researchers suggest means that throwing didn’t  develop as a means of hunting, but as a form of communication within  groups, i.e. throwing stuff at someone else became a form of self  expression, which is clearly evident to anyone who has ever been  targeted by a chimp locked up in a zoo.
(source, the same paper you mentioned, Researches find poop-throwing by chimps is a sign of intelligence)

And here you have the causes, in the same paper:
- It has been  hypothesized that neurological adaptations associated with evolutionary  selection for throwing may have served as a precursor for the emergence  of language and speech in early hominins.
-These results suggest  that chimpanzees that have learned to throw have developed greater  cortical connectivity between primary motor cortex and the Broca's area  homologue. It is suggested that during hominin evolution, after the  split between the lines leading to chimpanzees and humans, there was  intense selection on increased motor skills associated with throwing and  that this potentially formed the foundation for left hemisphere  specialization associated with language and speech found in modern  humans.

(source, the same paper you mentioned, Researches find poop-throwing by chimps is a sign of intelligence)

In a nutshell, the development of precisssion in poop throwing is associated with the cortical connectivity in brain structures essential for language in humans.

The easy joke is language came from the appropiate use of poop.
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But, if you ask me, the study goes very far in use of chimp natural behaviour to analyse early human development. As chimp can´t not use a very structurate group of sounds, due to morphology issues, for me there is a human biass in analyse their behaviour in this study.

domingo, 20 de noviembre de 2016

Is biology against equality?



 "Evolution selection of mammals are done by selecting the males. Such circumstances made polygamic male mammals significantly stronger and possibly smarter than their female counterparts.This is observed in chimps, lions etc"is this true?Because I'm not sure people who say that are against equality

 The question as it was posted.
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Someone requested me to anwer this bizarre question. Well, not so bizarre if you think on a female student whose teacher of biology is taking advance of his position to impose a distorted vision of the role of sex and gender among animals. Pure garbain. I really got upset when some stupid quoran told this anonyme question maker that they was being childish. 

My answer, go to learn about animals and you will discover that your rules are not universal and that the role of sex is far more complex that you thought. And fire that stupid teacher. 

Dicho de otro modo, que despidan a ese profesor, y que alguien mande a la mierda al gilipollas que insultó a quien quiera que hiciera esta pregunta. Creo que ese alguien voy a ser yo...mucha gente que pretende escribir en Quora dándoselas de estudiante, licenciado, etc...son gilipollas.
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A message for the anonyme op who a2aed me this:
I know you have unfollowed your question, and all I can do is to imagine what you had in mind. The quote has a grammal mistake, and also you could have explained your point more clearly, but I won't give it too many importance, not everybody here was educated in English, I wasn't educated in English, either. But I don't know wether you think the quote makes sense or not.
It doesn't.
If you are a young female student and have had to endure some stupid sexist insinuations in your biology tuitions, think on this, you are not the first one. Question your proffessor's intenttions. Stay strong and cold minded. And if you like biology, go on. It is incredibly varied and interesting, it has surprises for all of us, and it is experimenting a revolution in the way of regarding female mammals in the last years. 


 
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First off, Biology is not nature, it is  a science that studies a part of nature, life and living organisms branched in numerous subdisciplines that are defined by the scale at which those organisms are examined. Nature, on the other hand, is is the natural, physical, or material world or universe, it is life and the physical world at the same time. It is very important to make this distinction, which is in fact the cornerstone of this answer. You may not confuse our natural world with the science that studies it.
A science is a is a methodical system of acquiring knowledge, that seeks to explain the events of natural world in a reproductible way by other researhers working under the same conditions. In the case of Biology, the main stages of this method are observation and experimentation, and interpretation of the data acquired.
Although observation and experimentation are not free of flaws and assumptions, it is interpretation the most problematic part, because the uncoscious preconceptions, bias and assumptions afect it in a degree we can not predict.
Biology, biologists and description of the observational data made by biologists is not rid of assumptions, especially when it comes to matters that are important for our conception of the world, and the two main ones are the role of sexes (sadly, for some even of races) in human society and the connection of humans with the rest of animals. Put it in another words, you won't read more vivid discussions and more arrogant asumptions regarding biologic topics than in those two themes. Nobody will give so much thinking and energy to the way of feeding of mites or the reproduction of lichens. But when it comes to these two cathegories everybody seems to have something to say, no matter how much they really know about the science, the natural world, and all the incredibly intricated kind of complexities the study of animals, and mammals has.
I repeat, nobody is aware of how much their assumptions interfere the conclussions they make about the data proportioned by observations, and very few have take the job of looking for when did this theory of male rivalry conducting evolution started. It results that it started in the very writings of Darwin, who was assuming the preconceptions of the society he lived in:
Less widely known is that many evolutionists, including Darwin, taught that women were biologically and intellectually inferior to men. The intelligence gap that Darwinists believed existed between males and females was not minor, but of a level that caused some evolutionists to classify the sexes as two distinct psychological species, males as homo frontalis and females as homo parietalis. Darwin himself concluded that the differences between male and female humans were so enormous that he was amazed that ‘such different beings belong to the same species’ and he was surprised that ‘even greater differences still had not been evolved.’
Darwin concluded that males were like animal breeders, shaping women to their liking by sexual selection. In contrast, war pruned weaker men, allowing only the strong to come home and reproduce. Men were also the hunting specialists, an activity that pruned weaker men. Women by contrast, ‘specialized in the “gathering” part of the primitive economy.’
Sexism is not absent in the story of teaching of the evolution theory.
The female inferiority doctrine is an excellent example of the armchair logic, a logic derived from our assumptions imposed to the interpretation of data. Do you see that what Darwin theorized based upon his own preconceptions is exactly the core of the phrase quoted in the question?. Can you see it?.
Darwin taught that the differences between men and women were due partly, or even largely, to sexual selection. A male must prove himself physically and intellectually superior to other males in the competition for females to pass his genes on, whereas a woman must only be superior in sexual attraction. Darwin also concluded that ‘sexual selection depended on two different intraspecific activities: the male struggle with males for possession of females; and female choice of a mate.’
In Darwin’s words, evolution depended on ‘a struggle of individuals of one sex, generally males, for the possession of the other sex.’ (Darwin, C., The Origin of Species by Means of Natural Selection, 1897 edition, D. Appleton and Company, New York, p. 108, 1859.).
Those old victorian sexist prejudices absorbed in the understanding of biology of so many generations and appearing now, as of 2016, in this stupid phrase and in this interesting question. Memes. Dawkins had it right. Memes percolating in our collective unconscious.
Now that I have stablished the difference between nature and its study, the existence of unconscious bias in our way of understanding data acquired by observation of experiments, and he origin of assumption of that evolution is driven by selecting males...which is way much more than the former answers have done, I will take a time for a laugh. The rest of the answer is as painfully long and complex to explain as this part, so let's make a stop.


"[...] Such circumstances made polygamic male mammals significantly stronger and possibly smarter than their female counterparts.This is observed in chimps, lions etc". This is pissing, isn't it?. Absolutely!

Oh, you think that this is not fair...just cherrypicking?. What do you think that people propossing that selecion of males drives mammal evolution do?. Cherrypicking, no more. Worse, they can't give you a single study carried out on all poligamic mammal species that shows that evolution is made selecting males. NOT A SINGLE STUDY THAT BACKS THIS CONCLUSSION. Cope with it. There are not such studies.

Let's go again with the boring explanation.
"Evolution selection of mammals are done by selecting the males. Such circumstances made polygamic male mammals significantly stronger and possibly smarter than their female counterparts.This is observed in chimps, lions etc"
I'm going to be very clement here, and assume that the first phrase refers to polygamic mammals as well. It is more difficult to make a case against the phrase assuming that we are talking about polygynic species than in any other possibility, and I going to refer to this case. Also, I will forget about the pissing part of smarter males from now on.
To start off, we are talking about mammals. What all mammals have in common is that every single individual depends on their mother in the first part of their lives. We could think that those mammals produce equal number of male and female offspring, and that they invest the same work in rearing them up. But this is not always true.
Within species, mothers also often subtly adjust sex ratios to advantage themselves. Since the variance in male reproductive success is usually higher, it makes sense for mothers to produce more sons when they expect to have highly successful offspring, and more daughters when they don't. Boars, for instance, produce extra extra sons in small litters (when the mother can invest more) and extra daughters in large litters. Mother marsupials, including antechinus and brushtail possums, are thought to produce more sons to avoid competing with their daughters. Female bighorn sheep produce more sons when they are older and conditions are poor, but only breed every second season. The net outcome of this sort of individual-level bias is usually still a 50:50 sex ratio.
Here we examine the extent to which parental investment in red deer (Cervus elaphus) and other polygynous mammals matches these predictions. We conclude that, in several mammals, mothers invest more heavily in individual sons than daughters.
So we have here some evidences that point that parents make different choices accordingly to the possible reproductive success of their offspring accordingly to its sex. And they can favourite male over female or viceversa in function of the ecological factors. Parental selection of both sexes.
The matter is even more complicated, although infanticide commited by males due to sexual conflict is very estudied and divulgated (Infanticide (zoology), it is the known case of males killing the cubs when entering in a pride) there are other kinds of infanticide such as those commited by non maternal females due to sexual and resource competence (female rats will eat the kits of strange females for a source of nutrition, and to take over the nest for her own litter (wikipedia Infanticide (zoology). Mother rats also kill their own if they are deformed or wounded, to allow them to allocate resources to their other offspring. Why killing infants can benefit animals. Dominant females meerkats are known to kill subordinates' young, and they too have been known to kill a top female's young if they have a litter of their own. But male meerkats do not get blood on their paws. Why killing infants can benefit animals). Those killings don't select males, they kill all the youngs, if you follow my reasoning.
The pression of intraspecifical agression to youngs has lead to different strategies in order to protect the offsprings, one of them used by many female primates is paternity confusion where female primates mate with multiple partners so the males don't know who is the father. But they also have to have a look upon nearby females:
The adaptive function of copulation calls in female primates has been debated for years. One influential idea is that copulation calls are a sexually selected trait, which enables females to advertise their receptive state to males. Male-male competition ensues and females benefit by getting better mating partners and higher quality offspring. We analysed the copulation calling behaviour of wild female chimpanzees (Pan troglodytes schweinfurthii) at Budongo Forest, Uganda, but found no support for the male-male competition hypothesis. Hormone analysis showed that the calling behaviour of copulating females was unrelated to their fertile period and likelihood of conception. Instead, females called significantly more while with high-ranking males, but suppressed their calls if high-ranking females were nearby. Copulation calling may therefore be one potential strategy employed by female chimpanzees to advertise receptivity to high-ranked males, confuse paternity and secure future support from these socially important individuals. Competition between females can be dangerously high in wild chimpanzees, and our results indicate that females use their copulation calls strategically to minimise the risks associated with such competition.
Oops, female female competition in chimpanzees!.
With all these facts together, similar enviromental selective pressure, gestation and lactation periods, parental choices, female infaticide and female-female competition...the question is how can anybody say that selection is made only on males?. Is Higher male variance in reproductive success enough to contrarest all the above pressures and make the selective pressure on females insignificant?. There is only a way to make a case for it. A great study with the sufficient number of species and individuals to demonstrate it. And do you know what?. Nobody has done it.
So the believing in a greater selective pressure over males is just that, a believing. The theory was coined in the very first years of the evolution theory debate and it has never been properly demonstrated.
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There are only two exceptions in which the statement of a higher selective pressure over mammals males make sense:
- Harmful mutations on the X chormosome.
- Poultry and cattle: Where the asumption of "evolution done selecting the males" has some sense, is specifically in my speciality, agronomy. In fact I was also taught this same concept, but it is false. What it is true is that under non natural circumstances, where the animals can't make their own choices of reproduction, i.e. cattle and factory farm animals, it is more efficent to select the sperm of the males that have had very productive offspring (milky cows, meat cattle, egg laying chicks...), because you can very cheaply collect it, filter it in the way you can obtain many doses out of the original one, conserve it and sell it. This is the sense of this concept for an agronomist, and it makes sense. But it is not the real world under natural selection :).
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Final worlds about equality in humans and finally about nature:
So what? Equality of the sexes does not mean that women want to be as strong or as large as males on average! It means equality of rights and opportunities. Biology and evolution has nothing to do with human rights. A male lion that dethrones the king of the pride kills his cubs. It's what lions do: infanticide. If a bigger man kills a woman's husband so he can marry her and then kills her children, it's a crime. My point is, don't go invoking biology as justification for human behavior considered criminal, immoral or unacceptable. It's not going to help you much. Here's an undeniable fact of human biology: we are highly social animals with complex and changing culture.
Does nature take care about equality?. No. In fact there are terrible reproductive strategies for both males and females in nature.
Do we humans have to care about equality?. Yes, this concept is a product of our social evolution which is a product of our evolutive history, as it is the science of Biology. Nature is the whole picture, like the cosmos, biology is our way of making cosmology. (sorry, I needed to give some visual reference, I have a sort of visual brain).

viernes, 18 de noviembre de 2016

Why do so many huskies have heterochromia?

Very interesting question. It is obviously due to a evenly distribution of melanin between the two eyes.  In huskies it always includes one eye blue (1).

The reason is that this breed was developed by inbreeding together with the standards of the breed allowing blue eyes, and also heterochromia (for example malamute's standard is brown eyes, and heterochromia is not allowed), so the condition is not restricted by those standards.

Genetically, there are four ways of getting blue eyes in dogs, three of them related to the color of the coat causing white spots, patterns or diluted patterns, and only one that it is not linked to the coat. This last form is what huskies have and hence the color of the eye doesn't interefer with the normal pattern of the breed.
Breeders say that this is caused by a single gene, but i don't see they prove it in any way, because they don't tell wether it is a dominant character, codominant or recesive. So, i just tell what they say, but i don't give it entire credit.
This is the explanation given:

Lastly, blue eyes can be inherited as a completely separate gene,  unaffected by coat colour. This gene is, however, rare. It occurs  occasionally in the Border Collie and similar breeds, but mainly it's  seen in the Siberian Husky. Huskies can have one or both blue eyes, regardless of their main coat colour,  ranging in shade from almost white to sky blue. This is particularly striking when seen on black dogs.



Now, this only explains why do they inherit blue eyes independently of the pattern of the fur, but not why they have different eye color. And this last is very important, as they have the same genes in all their body, it seems that the development of the melanin is inhibited in one eye, and not in the other. This is not that strange (2). But what factor causes this inhibition is what remains unexplained, and I have found information regarding this issue.

Here is a good site about dog genetics:
Dog Coat Colour Genetics

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(1) Blue eyes are due to the Tyndall scattering of light in the eye, a phenomenon similar to that which accounts for the blueness of the sky called Rayleigh scattering. There are not blue pigments in the dogs eyes. Eye color is thus an instance of structural color and varies depending on the lighting conditions, especially for lighter-colored eyes.

(2) Heterochromia is very frequent in cats that have some form of albinism (there are various genes that lead to complete or partial inhibition of color in cats). This seems indicates that the "enviromental" factors opering during the development of both eyes are independent, although coat patterns tend to be very simmetrical.
I find this exciting to study the development of the fetus, and perhaps it could give us some ideas to understand some newborn defects such as cleft palates.
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As i told it is a very-very interesting quesion and I wish I knew more about this.